Eduardo Domingo Renoj-Matul, known as “Turko,” and his lieutenant, Cristobal Mejia-Chaj, were taken into custody on Friday and have pleaded not guilty to multiple charges related to migrant smuggling over a five-year period, according to the U.S. Attorney’s Office. A federal judge ordered both men, who are in the country illegally, to remain jailed without bond until their trial in April.

The indictment identifies Renoj-Matul as the leader of an extensive smuggling network that has been operating for at least 12 years, primarily transporting migrants from Guatemala to the U.S.

This is a developing news article and will be updated as more information is available.

Reporting by the Associated Press contributed to this article.

Two suspected leaders of a human smuggling ring accused of illegally bringing 20,000 people from Guatemala into the U.S. have been arrested in Los Angeles, federal prosecutors announced Monday.